Plecoptera Species File
species

Helopicus subvarians (Banks, 1920)

Vernal Springfly
Helopicus subvarians (Banks, 1920): Adult reared on 18 April from nymphs collected from Ausable River in southern Ontario 3/21/2010, M. Pessino collector. (Otu).
Helopicus subvarians (Banks, 1920): Adult reared on 18 April from nymphs collected from Ausable River in southern Ontario 3/21/2010, M. Pessino collector. (Otu).
Helopicus subvarians (Banks, 1920): Helopicus subvarians female reared 18 April from nymph collected 21 March 2010 from the Ausable River, southern Ontario, M. Pessino collector. (Otu).
Helopicus subvarians (Banks, 1920): Nymph collected 3/21/2010 from Ausable River, southern Ontario by M. Pessino. (Otu).

Nomenclature (27)

  • Perla subvarians Banks, 1920: 317.

    Syntype; MCZ M. C. Z. 10,817; deposited at: Museum of Comparative Zoology (MCZ); United States: Great Falls. Syntype; MCZ M. C. Z. 10,817'; deposited at: Museum of Comparative Zoology (MCZ); United States: Maryland: Great Falls, High Island

  • Perla tincta Needham & Claassen, 1925: 89. (homonym)

    Holotype; female; c1354d45-655f-4a49-b454-08d32978c9f4; deposited at: Cornell University Insect Collection (CUIC); United States: North Carolina

  • ... Show all ... (23)
  • Helopicus subvarians (Banks, 1920) in Hogan & Grubbs, 2022: 400.
  • Perla subvarians Banks, 1920 in Myers, Kondratieff, Grubbs, Pett, DeWalt, Mihuc & Hart, 2025: 71.

Nomenclature references (22)

  • Banks, N. (1920) New Neuropteroid Insects. Bulletin of the Museum of Comparative Zoology at Harvard College, 64(3), 299–362, 7 pls. Available at http://www.biodiversitylibrary.org/item/28057#page/307/mode/1up
  • Claassen, P.W. (1936) New Names for Stoneflies (Plecoptera). Annals of the Entomological Society of America, 29(4), 622–623. Available at http://chla.library.cornell.edu/c/chla/browse/title/5077679.html#1936
  • ... Show all ... (18)
  • Claassen, P.W. (1940) A catalogue of the Plecoptera of the world. Memoirs of the Cornell University Agricultural Experiment Station, 232, 1–235. Available at https://reader.library.cornell.edu/docviewer/digital?id=chla7251474_8571_012#mode/1up
  • Giberson, D.J. & Garnett, H.L. (1996) Species composition, distribution, and summer emergence phenology of stoneflies (Insecta: Plecoptera) from Catamaran Brook, New Brunswick. Canadian Journal of Zoology, 74, 1260–1267.
  • Grubbs, S. (2011) A review of stonefly (Insecta; Plecoptera) taxonomic research in Alabama, with new state records and an updated checklist. Illiesia: International Journal of Stonefly Research, 7(2), 24–30. Available at http://illiesia.speciesfile.org/papers/Illiesia07-02.pdf
  • Grubbs, S.A. & Sheldon, A.L. (2018) The stoneflies (Insecta, Plecoptera) of the Talladega Mountain region, Alabama, USA: distribution, elevation, endemism, and rarity patterns. Biodiversity Data Journal, 6(e22839), 1–40. Available at http://doi.org/10.3897/BDJ.6.e22839
  • Hogan, P.N. & Grubbs, S.A. (2022) Distributional Trends and Species Richness of Maryland, USA, Stoneflies (Insecta: Plecoptera), with an Emphasis on the Appalachian Region. Ecologies, 3(3), 395–421. https://doi.org/10.3390/ecologies3030029
  • Illies, J. (1966) Katalog der rezenten Plecoptera. Das Tierreich – Eine Zusammenstellung und Kennzeichnung der rezenten Tierformen., 82, 1–631.
  • Kondratieff, B.C. & Kirchner, R.F. (1987) Additions, taxonomic corrections, and faunal affinities of the stoneflies (Plecoptera) of Virginia, USA. Proceedings of the Entomological Society of Washington, 89(1), 24–30. Available at http://https://www.biodiversitylibrary.org/page/16262424
  • Kondratieff, B.C. & Voshell, J.R., Jr. (1979) Checklist of the stoneflies (Plecoptera) of Virginia. Entomological News, 90(5), 241–246.
  • Kondratieff, B.C. (2004) Perlodidae-Perlodinae (The Springflies). In Ohio Biological Survey Bulletin New Series. Vol. 14, Issue 4, pp. 149–180.
  • Kondratieff, B.C., Verdone, C.J. & Roble, S.M. (2017) New records of stoneflies (Plecoptera) from Virginia, U.S.A. Perla, 35, 22–27. Available at http://plecoptera.speciesfile.org/HomePage/Plecoptera/LitArchive/PerlaNo35_2017.pdf
  • Myers, L., Kondratieff, B.C., Mihuc, T.B. & Ruiter, D.E. (2011) The Mayflies (Ephemeroptera), Stoneflies (Plecoptera), and Caddisflies (Trichoptera) of the Adirondack Park (New York State). Transactions of the American Entomological Society, 137(1+2), 63–140. Available at http://https://doi.org/10.3157/061.137.0118PY
  • Myers, L.W., Kondratieff, B.C., Grubbs, S.A., Pett, L.A., DeWalt, R.E., Mihuc, T.B. & Hart, L.V. (2025) Distributional and species richness patterns of the stoneflies (Insecta, Plecoptera) in New York State. Biodiversity Data Journal, 13(e158952), 1–115. https://doi.org/10.3897/bdj.13.e158952
  • Needham, J.G. & Claassen, P.W. (1925) A Monograph of the Plecoptera or Stoneflies of America North of Mexico. The Thomas Say Foundation, II, 1–397. Available at http://www.archive.org/details/monographofpleco00need
  • Pescador, M.L., Rasmussen, A.K. & Richard, B.A. (2000) A Guide to the Stoneflies (Plecoptera) of Florida. State of Florida, Department of Environmental Protection, Division of Water Resource Management, Tallahasee, Final Report for DEP Contract Number WM671, 94 pp. & Appendices.
  • Ricker, W.E. (1952) Systematic Studies in Plecoptera. Indiana University Publications Science Series, 18, 1–200.
  • Stark, B.P. & Ray, D.H. (1983) A revision of the genus Helopicus (Plecoptera: Perlodidae). Freshwater Invertebrate Biology, 2(1), 16–27.
  • Tarter, D.C. & Kirchner, R.F. (1980) List of the Stoneflies (Plecoptera) of West Virginia. Entomological News, 91(2), 49–53. Available at http://www.biodiversitylibrary.org/page/2693425
  • Tarter, D.C. & Nelson, C.H. (2010) New state, county, and drainage basin records of West Virginia (USA) stoneflies (Plecoptera). Entomological News, 121(2), 159–162. Available at http://www.bioone.org/doi/full/10.3157/021.121.0206
  • Tarter, D.C., Chaffee, D.L. & Grubbs, S. (2006) Revised checklist of the stoneflies (Plecoptera) of Kentucky, U.S.A. Entomological News, 117(1), 1–10.
  • Zwick, P. (1973) Insecta: Plecoptera Phylogenetisches System und Katalog. Das Tierreich – Eine Zusammenstellung und Kennzeichnung der rezenten Tierformen., 94, 465 pp.

Descendants and synonyms

Stats

Summary of taxonomic names by rank, showing valid extant names, fossil names, invalid names, and total counts.
Taxon Valid names
Rank Extant Fossil Invalid Total
species1045